Monophyletic, but nested within Oreogrammitis/Radiogrammitis (Sundue et al., 2014). Themelium is supported as monophyletic in Sundue et al. (2014), but nested within a clade containing polyphyletic Oreogrammitis and Radiogrammitis. The types of Oreogrammitis and Themelium remain to be sampled, and thus we recommend maintaining these pending further research.
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